QTLdb: entrichment tests

Release 58
(Dec 29, 2025)

Whole genome analysis for QTL/association enrichment

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Version: Enrich S: beta v0.8

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Data:

Number of milk composition - fat traits:	73
Number of QTL / associations found:	47,401
Number of chromosomes where QTL / associations are found:	30

Chi-squared (χ²) test: are milk composition - fat traits over-represented on some chromosomes?

Chromosomes	Total χ²	df	p-values	FDR ^*	Size of χ²
Chromosome X	92248.94879	29	9e-41	9.310345e-41
Chromosome 1	68657.46652	29	9e-41	9.310345e-41
Chromosome 2	62279.62515	29	9e-41	9.310345e-41
Chromosome 3	80383.72954	29	9e-41	9.310345e-41
Chromosome 4	91727.41270	29	9e-41	9.310345e-41
Chromosome 5	556297.63880	29	9e-41	9.310345e-41
Chromosome 6	37094.10904	29	9e-41	9.310345e-41
Chromosome 7	77485.15504	29	9e-41	9.310345e-41
Chromosome 8	82963.64298	29	9e-41	9.310345e-41
Chromosome 9	69902.12382	29	9e-41	9.310345e-41
Chromosome 10	66867.01450	29	9e-41	9.310345e-41
Chromosome 11	29271.59402	29	9e-41	9.310345e-41
Chromosome 12	84581.15362	29	9e-41	9.310345e-41
Chromosome 13	13126.99578	29	9e-41	9.310345e-41
Chromosome 14	6930819.71997	29	9e-41	9.310345e-41
Chromosome 15	48354.42036	29	9e-41	9.310345e-41
Chromosome 16	40566.42120	29	9e-41	9.310345e-41
Chromosome 17	3344.01977	29	9e-41	9.310345e-41
Chromosome 18	89784.82211	29	9e-41	9.310345e-41
Chromosome 19	161.00953	29	3.025553e-20	3.025553e-20
Chromosome 20	1464.39825	29	2.123527e-290	6.370581e-289
Chromosome 21	93034.02364	29	9e-41	9.310345e-41
Chromosome 22	99028.12153	29	9e-41	9.310345e-41
Chromosome 23	87100.12374	29	9e-41	9.310345e-41
Chromosome 24	100658.14948	29	9e-41	9.310345e-41
Chromosome 25	1039.99742	29	3.02966e-200	4.544490e-199
Chromosome 26	5871283.57330	29	9e-41	9.310345e-41
Chromosome 27	27823.85875	29	9e-41	9.310345e-41
Chromosome 28	100249.39474	29	9e-41	9.310345e-41
Chromosome 29	93690.79442	29	9e-41	9.310345e-41

Chi-squared (χ²) test: Which of the 73 milk composition - fat traits are over-represented in the QTLdb

Traits	Total χ²	df	p-values	FDR ^*	Size of χ²
Milk C10 index	80.63798	72	0.2272410	2.591968e-01
Milk C14 index	10164.27619	72	9e-41	2.119355e-40
Milk C16 index	2807.78495	72	9e-41	2.119355e-40
Milk C17 index	670.59163	72	6.402378e-98	7.789560e-97
Milk C18 index	385.76254	72	1.949878e-44	9.489406e-44
Milk arachidic acid content	182.36304	72	1.544005e-11	2.348174e-11
Milk arachidonic acid content	357.56601	72	1.847606e-39	4.214851e-39
Milk beta-carotene content	367.92623	72	2.806446e-41	1.280441e-40
Milk branched-chain fatty acid content	86.75854	72	0.1132635	1.312418e-01
Milk butyric acid content	17706.2529	72	9e-41	2.119355e-40
Milk capric acid content	3163.14458	72	9e-41	2.119355e-40
Milk caproic acid content	5042.25746	72	9e-41	2.119355e-40
Milk caprylic acid content	2787.42913	72	9e-41	2.119355e-40
Milk cis-12-Octadecenoic acid content	170.93838	72	5.052588e-10	7.376778e-10
Milk cis-4,7,10,13,16,19-Docosahexaenoic acid content	253.07714	72	5.5902e-22	9.953283e-22
Milk cis-7,10,13,16,19-Docosapentaenoic acid content	78.92751	72	0.2693061	3.024515e-01
Milk cis-8,11,14-Eicosatrienoic acid content	530.39215	72	4.995843e-71	3.670163e-70
Milk cis-9,trans-11,cis-15-Octadecatrienoic acid content	30.16437	72	0.9999966	1.000000e+00
Milk cis-9-C10:1 fatty acid content	136.86558	72	6.295831e-06	8.671616e-06
Milk cis-9-Eicosenoic acid content	16.74877	72	1	1.000000e+00
Milk cis-9-Heptadecenoic acid content	41.66407	72	0.9984136	1.000000e+00
Milk cis-fatty acid content	110.55014	72	0.002370896	2.884590e-03
Milk cis-vaccenic acid content	216.90268	72	1.930648e-16	3.277612e-16
Milk conjugated linoleic acid content	556.33864	72	6.128236e-76	5.592015e-75
Milk docosanoic acid content	88.65291	72	0.08886066	1.046263e-01
Milk eicosapentaenoic acid content	512.44663	72	1.187213e-67	7.878777e-67
Milk fat content	300.88757	72	9.350824e-30	1.896139e-29
Milk fat percentage	10263.19987	72	9e-41	2.119355e-40
Milk fat yield	6033.73855	72	9e-41	2.119355e-40
Milk fat-to-protein ratio	95.61497	72	0.03283939	3.929960e-02
Milk fatty acid unsaturated index	530.37755	72	5.027621e-71	3.670163e-70
Milk heptadecenoic acid content	423.08673	72	3.808006e-51	1.985603e-50
Milk lauric acid content	3110.42157	72	9e-41	2.119355e-40
Milk lauroleic acid content	71.68239	72	0.48839	5.321264e-01
Milk linoleic acid content	1799.60098	72	9e-41	2.119355e-40
Milk linolenic acid content	1648.15985	72	1.479633e-296	1.080132e-294
Milk long-chain fatty acid content	142.21881	72	1.601868e-06	2.292870e-06
Milk margaric acid content	180.68399	72	2.600386e-11	3.874044e-11
Milk medium-chain fatty acid content	726.72647	72	6.837547e-109	1.247852e-107
Milk monounsaturated fatty acid content	269.11962	72	1.543698e-24	2.889486e-24
Milk myristelaidic acid content	302.85258	72	4.387978e-30	9.152068e-30
Milk myristic acid content	6200.21029	72	9e-41	2.119355e-40
Milk myristoleic acid content	8059.80558	72	9e-41	2.119355e-40
Milk octadecenoic acid content	302.85258	72	4.387978e-30	9.152068e-30
Milk odd-chain fatty acid percentage	30.16437	72	0.9999966	1.000000e+00
Milk oleic acid content	1226.68342	72	1.621464e-209	3.945562e-208
Milk omega-6 fatty acid content	207.46623	72	4.654076e-15	7.385816e-15
Milk palmitelaidic acid content	2.42791	72	1	1.000000e+00
Milk palmitic acid content	7165.31191	72	9e-41	2.119355e-40
Milk palmitoleic acid content	1953.56937	72	9e-41	2.119355e-40
Milk pentadecylic acid content	565.30514	72	1.208741e-77	1.260544e-76
Milk polyunsaturated fatty acid content	133.3217	72	1.516416e-05	2.049970e-05
Milk rumenic acid content	188.8296	72	2.020283e-12	3.137886e-12
Milk saturated fatty acid content	261.26028	72	2.813014e-23	5.133751e-23
Milk saturated to unsaturated fatty acid ratio	210.58932	72	1.635458e-15	2.653076e-15
Milk short-chain fatty acid content	130.95208	72	2.695228e-05	3.577303e-05
Milk stearic acid content	2605.54223	72	9e-41	2.119355e-40
Milk tetracosanoic acid content	674.92141	72	9.197017e-99	1.342764e-97
Milk trans-10,cis-12-octadecadienoic acid content	302.85258	72	4.387978e-30	9.152068e-30
Milk trans-10-Octadecenoic acid content	137.13483	72	5.883786e-06	8.259930e-06
Milk trans-11,cis-15-Octadecadienoic acid content	276.19884	72	1.102071e-25	2.117136e-25
Milk trans-16-Octadecenoic acid content	232.78651	72	7.896216e-19	1.372438e-18
Milk trans-4-Octadecenoic acid content	30.16437	72	0.9999966	1.000000e+00
Milk trans-6/8-Octadecenoic acid content	72.64264	72	0.4566227	5.050524e-01
Milk trans-9,trans-12-Octadecadienoic acid content	478.46685	72	2.600824e-61	1.582168e-60
Milk trans-9-Octadecenoic acid content	471.14809	72	5.90472e-60	3.315727e-59
Milk trans-fatty acid content	118.98128	72	0.0004162225	5.149872e-04
Milk trans-fatty acid to cis-fatty acid ratio	122.16945	72	0.0002066753	2.601258e-04
Milk trans-vaccenic acid content	130.35787	72	3.108275e-05	3.980773e-05
Milk tricosanoic acid content	283.93207	72	6.008986e-27	1.185557e-26
Milk tridecylic acid content	1439.48151	72	2.686025e-253	9.803991e-252
Milk undecylic acid content	213.3446	72	6.460248e-16	1.071814e-15
Milk unsaturated fatty acid content	130.43055	72	3.054646e-05	3.980773e-05

Correlations found between some of these traits for your reference

No correlation data found on these traits

Overall Test

Data		Chi'Square Test			Fisher's Exact Test
Number of chrom.:	30	χ²	=	15001289.458510
Number of traits:	73	df	=	2088
Number of QTLs:	47,401	p-value	=	0

FOOT NOTE: * : FDR is short for "false discovery rate", representing the expected proportion of type I errors. A type I error is where you incorrectly reject the null hypothesis, i.e. you get a false positive. It's statistical definition is FDR = E(V/R | R > 0) P(R > 0), where V = Number of Type I errors (false positives); R = Number of rejected hypotheses. Benjamini–Hochberg procedure is a practical way to estimate FDR.