NUTRITION                                         PIH-3

PURDUE UNIVERSITY.  COOPERATIVE EXTENSION SERVICE.
WEST LAFAYETTE, INDIANA



                      Dietary Energy for Swine

Authors
Palmer J. Holden, Iowa State University
Gerald C. Shurson, University of Minnesota
James E. Pettigrew, University of Minnesota

Reviewers
Richard C. Ewan, Iowa State University
John P. Hitchcock, University of Tennessee
William G. Luce, Oklahoma State University



     Pigs require energy to maintain normal  body  processes,  to
grow  and  to  reproduce.  Feeds  supplying energy are major com-
ponents of all swine diets, and the quantity of diet  voluntarily
consumed  by pigs is related to its energy content. Carbohydrates
from cereal grains are the most abundant energy source  in  swine
diets.  Fats  and oils contain more energy than carbohydrates per
unit weight but are included to a lesser extent. Amino acids,  or
protein,  may  serve as an energy source if included in the diets
in excess of animals' requirement for protein synthesis.

     The value of a feedstuff is based on several factors:  pala-
tability  (how  well the material will be consumed by an animal),
availability of energy  and  the  feed's  contribution  of  other
nutrients  (protein or amino acids, vitamins, minerals). Should a
pork producer buy corn, wheat, or oats as a feed ingredient? This
depends  primarily  on  the  cost  of these ingredients and their
value as sources of energy and other nutrients for the pig.


Measurement of Energy

     To make sound decisions in selecting feed ingredients, it is
desirable  to  have  an  understanding  of  the  system  by which
feedstuffs are rated for energy content and the use of these rat-
ings  toward meeting the energy requirements for pigs' growth and
production. The gross energy (GE) of a feed ingredient is defined
as the heat produced when a substance is burned.  It is expressed
as calories per unit weight. A calorie  is  the  amount  of  heat
required  to raise the temperature of one gram of water from 14.5
to 15.5 degrees C. A kilocalorie (kcal) is 1,000 calories, and  a
megacalorie (mcal) is a million calories.

     Not all of the feed consumed is digested and absorbed.  Some
energy is lost in the fecal material (Fig. 1). Thus, GE is a poor
estimate of energy for the pig. The amount  of  energy  remaining
after  subtracting the fecal energy loss from total energy intake
is designated as digestible energy (DE). The  difference  between
GE  and DE may be large.  The greater the digestibility of energy
(DE/GE) the greater its value  as  a  source  of  energy  to  the
animal.  DE  is a more meaningful measure for livestock producers
than GE.

     Metabolizable energy (ME) is the "usable" energy of  a  feed
for  the  pig to live and grow and is obtained by subtracting the
urinary energy loss from the DE.  In  most  cases,  metabolizable
energy represents approximately 95% of the digestible energy con-
tent, so the conversion from DE to ME can be made easily.

     Some energy is released as heat as a result  of  inefficien-
cies  in the metabolism of the nutrients. This is called the heat
increment (HI). It can be used only to keep the animal warm; heat
increment  produced  beyond that needed to maintain body tempera-
ture is wasted. The remaining energy is called  net  energy  (NE)
and  is used for maintenance (NEm) and production (NEp). Determi-
nation of NE values  requires  special  equipment  and/or  animal
feeding trials.


Major Energy Sources

Cereal Grains

     The basic energy sources for swine are  the  cereal  grains:
corn,  sorghum  grain  (milo),  barley, wheat, oats and their by-
products. Cereal  grains  are  high  in  carbohydrates  (starch),
palatable and highly digestible. Usually they contain less lysine
and other amino acids, minerals and vitamins than swine  require.
The  diets  therefore  must  be  supplemented with other feeds to
increase these nutrients to recommended levels.

     Grain by-products have many characteristics of their  origi-
nal  source  but  tend  to be bulkier and have less metabolizable
energy. Although their protein content usually is increased,  the
protein quality often is rather poor.

     Corn contains less protein but more energy  than  the  other
cereals.  Like all cereals, the composition of corn is influenced
by variety, growth conditions, method of harvesting and  storage.
Because  of  its  abundance and readily available energy, corn is
used as the base grain for comparing the nutritive value of other
cereal  grains.  Grinding is recommended except for high moisture
corn.

     Sorghum grain is similar in quality to  corn  and  can  com-
pletely  replace  corn  in swine diets. Its energy value is about
95% of the value of corn except for some bird-resistant varieties
which  may  be  only 80% to 90% of the value of corn. Grinding is
recommended because the grain is rather small and hard.

     Barley contains more protein and fiber than corn. High qual-
ity  barley  has 90% to 100% of the feeding value of corn, but it
may be less palatable.

     Wheat is equal to corn in feeding value and is  very  palat-
able  when  it  is medium to coarsely ground in complete rations.
Wheat can completely replace corn in swine diets.

     Oats contain more protein than  corn,  but  their  value  in
swine diets is only 90% of corn because of their higher fiber and
lower energy content.

Fats and Oils

     Fats and oils contain about 2.25 times as much metabolizable
energy  per  pound as cereal grains, but they are more expensive.
Fats are available commercially in products  such  as  bleachable
fancy tallow, prime tallow, yellow grease, hydrogenated vegetable
fat and various dry fat products which include fat and a dry car-
rier.  Fat  sources  should  be  protected  from  rancidity by an
antioxidant. There do not  appear  to  be  major  differences  in
dietary  value  among the fat sources, except for very young pigs
which may not utilize the harder animal fats as  well  as  softer
vegetable  fats.  Vegetable oils and the dry fat products tend to
be more expensive than animal fats.

     Supplemental fat is  difficult  to  mix  in  on-farm  mixing
facilities,  especially in cold weather. Fat changes the physical
characteristics of a swine feed.  Feed containing  added  fat  is
somewhat  sticky  and  therefore tends to bridge in bulk bins and
feeders. It tends to "oil out" of paper bags  and  reduce  pellet
hardness.  These problems increase as the fat level increases and
become severe when the added fat exceeds 6% of  the  feed.  Added
fat  reduces  dustiness and therefore improves the environment in
swine buildings.

     Certain biological effects also can be expected when fat  is
added  to  diets  of  starting, growing and finishing pigs. These
include: improved palatability, reduced feed consumption,  signi-
ficant  improvement in feed efficiency, slight increase in growth
rate, increased carcass fatness at high fat levels and high  lev-
els of vegetable oils causing a softening of carcass fat.

     Pigs' responses to  fat  may  be  greater  in  warm  or  hot
environments  than  in  cool  environments.  Fat has a lower heat
increment than carbohydrates and proteins and is less  likely  to
cause reduced feed intake during heat stress.

     When fat is added to a swine diet, the amount of  feed  con-
sumed  usually  decreases.  However,  animals'  needs  for  other
nutrients remain relatively constant when expressed  on  a  daily
basis.  Therefore,  to  maintain performance when fat is added to
the  diet,  the  concentration  of  other  nutrients  should   be
increased.

     The decision to add fat is an economic one.  If  improvement
in growing-finishing swine preformance more than offsets the cost
of adding fat, it is economical. Typically, adding 1% fat to  the
diet results in approximately 2% better efficiency.

     When the piglet survival rate is  below  85%,  supplementing
the  sows'  diet  with fat during late gestation may improve sur-
vival rate. The added fat must provide at least 2.0  lb.  to  2.5
lb.  of  fat to each sow prior to farrowing. This appears to be a
response to fat and not to increased energy intake. The added fat
increases  the  fat  content  of  the colostrum and milk which is
responsible for the increased survival rate. Adding fat to lacta-
tion  diets increases voluntary ME consumption, but only slightly
reduces the weight loss in the sows. It increases weaning weights
of the litters due to increased fat in the milk. There is no evi-
dence that added fat improves subsequent reproductive preformance
of sows.


Fiber Content

     Some energy sources are relatively high in fiber and  reduce
gain  and  efficiency if fed at excessive levels. Pigs 40 lb. and
heavier usually can tolerate up to 5% of a high-fiber  feed  such
as  alfalfa  in their diet without a noticeable effect on perfor-
mance. As pigs mature, more and more low energy-high fiber  feeds
can be fed, especially to sows during gestation and post-weaning.
Fiber feeds such as wheat bran and beet pulp  may  be  useful  in
gestation  and farrowing diets because of their laxative effects,
but should constitute no more than 5% of a lactation diet.

     Fiber has a high heat increment and during cold stress  this
heat  can be utilized to maintain body temperature, thus, less is
wasted. Therefore, there is  a  smaller  difference  in  relative
values  between  fibrous grains (such as barley or oats) and corn
in cold weather. Conversely, in hot weather the high heat  incre-
ment becomes a problem for the pigs' cooling ability.


Moisture Content

     High-moisture grains contain less energy per pound  of  feed
because  of the water content. More pounds of high-moisture grain
must be used to get the same amount of dry matter.  Studies  with
high-moisture  grains fed in complete diets indicate similar per-
formance to dried grain when efficiency  is  measured  on  a  dry
matter  basis.  Free-choice feeding of grain and supplement often
results in poorer efficiency. See PIH-73,  High  Moisture  Grains
for Swine.


Grinding

     With the exception of high-moisture corn, grinding  improves
feeding  efficiency  for all grains, especially high-fiber grains
such as  oats  or  barley.  Finer  grinding  usually  results  in
improved  efficiency,  although  finely ground corn increases the
incidence of ulcers in finishing swine.  Fine  grinding  is  most
advantageous  for  pigs under 40 lb. Wheat is very palatable when
it is medium to coarsely ground in complete  rations  since  high
levels  of  finely ground wheat in diets has been associated with
lowered palatability due to pastiness of the  meal.  See  PIH-71,
Physical Forms of Feed.


Pelleting

     Pelleting a diet may increase gains by  5%  and  feed  effi-
ciency by 5% to 10%. A high-energy cereal such as corn or sorghum
benefits less from pelleting than fibrous feeds  like  barley  or
oats.  When  a  complete diet is purchased, pelleted diets may be
more economical than meal diets. However, the advantage  of  pel-
leting  probably  will  not offset the cost of hauling grain from
the farm to a pelleter and home again.


Relative Value

     In selecting energy sources for swine diets,  also  consider
protein  quality  and  content.  Because  the amino acids lysine,
tryptophan, threonine and methionine can  be  limiting  in  swine
diets,  levels of these amino acids on cereal grains affect their
overall value. Although sugar, molasses  and  fats  or  oils  are
energy sources, they provide little or no protein to the diets.

     The amount of feed per unit of gain is not the  most  impor-
tant  factor  in  swine  nutrition. Cost per unit of gain is more
important; therefore, it is necessary to use the most  economical
feed  sources  available  in  swine  diets.  The relative feeding
values shown in Tables 1 and 2 can be  used  to  determine  which
ingredient  is  most  economical.  For example, if corn costs 5.0
cents per pound, barley is worth about 4.5 cents per  pound  (5.0
cents  x  90%). If barley can be purchased for less than this, it
is a better buy.

______________________________________________________________________

Table 1. Relative feeding values of energy sources1.
_________________________________________________________________
                              Metabo- Relative  Maximum recommended 
                              lizable  feeding  percent of complete 
                                                diets3
                              energy   value vs.  Gesta-  Lacta-        
Ingredient (air dry)          kcal/lb. corn,%  2   tion     tion
__________________________________________________________________
Alfalfa meal dehydrated        775      70-80       50       5
Alfalfa meal sun-cured         800      60-70       50       5    
Animal fat stabilized         3585     185-210      10       6   
Bakery surplus material       1650      95-110      40      40   
Barley (48 lb/bu)             1380      90-100      80      80
Beet pulp, dried              1020      70-80       10       5 
Brewers dried grains          1000      90-100      40       5
Buckwheat                     1200      80-90       50       0
Corn yellow                   1550         100      80      80
Corn (high lysine)            1520     100-105      90      90  
Corn and cob meal             1300      80-90       70       0
Corn distillers dried grains  1540     115-130      40      10  
 with solubles
Corn gluten feed              1285         74       90       5 
Corn grits by-product (hominy)1400     100-105      60      60  
Emmer                         1140      80-90       20       0 
Sorghum grain (milo)4         1480      95-100(4)   80      80 
Millet (Proso)                1230      90-95       80      80
Molasses (77% D.M.)           1060      55-65        5       5
Oats (36 lb/bu)               1240      85-95       80       5
Oat groats                    1550     115-125       0       0
Potatoes (22% D.M.)            370      20-25       80       0
Rice grain                    1070         75       40       0  
Rye                           1300         90       20       0 
Spelt                         1180         85       40       0
Sugar                         1383      70-80        0       0
Triticale                     1450      90-95       80      80
Wheat, hard                   1475     100-105      80      80
Wheat, soft                   1500      90-95       80      80
Wheat, high protein           1500     100-105      80      80
Wheat bran                     980      60-65       30       5
Wheat middlings               1340     125-140      30       5
Wheat, dried                  1405     135-145       5       5
__________________________________________________________________


Contd ...Table 1.
_________________________________________________________________
                                Maximum recommended 
                                percent of complete diet3
                                                
                        Starter  Grow-   Remarks 
Ingredient (air dry)            finish
__________________________________________________________________

Alfalfa meal dehydrated       0    5    Good source of B vitamins
Alfalfa meal sun-cured        0    5    Unpalatable to baby pigs
Animal fat stabilized         5   10    High energy, reduces dust
Bakery surplus material      20   40    High energy, variable salt 
                                           content
Barley (48 lb/bu)            25   85    Corn substitute, lower energy
Beet pulp, dried              0    0    Bulky, high fiber, laxative
Brewers dried grains          0   10    High fiber, B-vitamin source,
                                          low lysine
Buckwheat                     0   50    Not to light colored pigs 
                                          outdoors
Corn yellow                  60   85    High energy, low lysine
Corn (high lysine)           60   90    Lysine analysis recommended
Corn and cob meal             0    0    Bulky, low energy
Corn distillers dried grains  5   10    B-vitamin source, low lysine
 with solubles
Corn gluten feed              5   25    Low lysine
Corn grits by-product (hominy)0   60    Subject to rancidity
Emmer                         0    0 
Sorghum grain (milo)4        60   85    Low lysine
Millet (Proso)               60   85    Low lysine
Molasses (77% D.M.)           5    5    Low energy, partial grain 
                                           substitute
Oats (36 lb/bu)               0   20    Low energy, partial grain 
                                           substitute
Oat groats                   20    0    Palatable
Potatoes (22% D.M.)           0   30    Must be cooked, low protein
Rice grain                    0   20    Low energy, low lysine
Rye                           0   25    Possible ergot toxicity, low 
                                           palatability
Spelt                         0   25    Low energy, low lysine
Sugar                         5    0    High palatability, no protein
Triticale                    20   85    Possible ergot
Wheat, hard                  60   85    Low lysine
Wheat, soft                  60   85    Low lysine
Wheat, high protein          60   85    Low lysine
Wheat bran                    0    0    Bulky, high fiber, laxative
Wheat middlings               0   10    Partial grain substitute
Wheat, dried                 20    5    High lactose content, variable 
                                           content
____________________________________________________________________

1Based on an air dry basis unless otherwise noted. High  moisture
feedstuffs  must  be converted to an air dry equivalent of 88-90%
dry matter to determine energy and substitution  rates.  Complete
data on all ingredients are not available.

2When fed at no more  than  maximum  recommended  %  of  complete
diets.  Relative values based on metabolizable energy, lysine and
phosphorus content using simultaneous equations. Example:

     ME    Lysine   Phosphorus   Price
___________________________________________________________
1550X +   .024Y +      0.25Z =   $/cwt. corn
1460X +   2.90Y +      0.60Z =   $/cwt. soybean meal (44%)
   0X +      0Y +     18.50Z =   $/cwt. dicalcium phosphate
Determine values for X, Y, and Z and multiply them times the M.E.
(kcal/lb), % lysine, and % phorphorus of feed in question and sum
the values.
3Higher levels may be fed although performance may decrease.
4Some bird resistant sorghums are 80-90% vs. corn.
______________________________________________________________________


______________________________________________________________________

Table 2. Relative feeding values of protein sources1.
_____________________________________________________________________
                        Metabo-    Relative  Maximum recommended
                        lizable     feeding  percent of complete
                                               diets3
                         energy    value vs.      Gesta-    Lacta-   
Ingredient (air dry)     kcal/lb. 44% soybean,%2   tion     tion  
_____________________________________________________________________
Blood meal, flash dried    1200      185-200        5        0  
Buttermilk, dry            1400       75-85         0        5  
Canola meal (solvent)      1200       75-85         5        5  
Corn gluten meal           1395       40-60         5        5  
Cottonseed meal, solvent   1150       65-75         5        5  
Distillers dried solubles  1180       65-70         5        5  
(corn)
Feather meal, hydrolyzed   1000       60-70         3        3  
Fish meal, anchovy         1120      140-165        5        5  
Fish meal, menhaden        1500      140-165        5        5  
Fish solubles (50% solids)  780       50-60         3        3  
Linseed meal               1280       55-65         5        5  
Meat and bone meal         1035       95-110       10        5  
                                                            
Peanut meal, expeller      1400       70-80         5        5  
Skim milk, dried           1620       95-100        0        0  
Sorghum gluten meal        1460       40-55         5        5  
Soybeans, whole, cooked    1640       90-100       30       25  
                           
Soybean meal, solvent      1460          100       25       20  
Soybean meal, solvent,     1535      110-112       22       18  
dehulled
Tankage (meat meal)         980      115-130       10        5  
Yeast, brewers dried       1400      100-105        3        3   
______________________________________________________________________


Contd ..Table 2.
______________________________________________________________________
                                Maximum recommended 
                                percent of complete diet3
                                                
                        Starter  Grow-   Remarks 
Ingredient (air dry)            finish
______________________________________________________________________
Blood meal, flash dried     5      5   Low isoleucine, unpalatable
Buttermilk, dry            20      5   Good amino acid balance
Canola meal (solvent)       5     10   Toxic problem at high levels
Corn gluten meal            0      5   Low lysine
Cottonseed meal, solvent    0      5   Gossypol toxicity, low lysine
Distillers dried solubles   5      5   B-vitamin source, low lysine
(corn)
Feather meal, hydrolyzed    0      3   Low lysine
Fish meal, anchovy          5      5   Excellent amino acid balance
Fish meal, menhaden         5      5   Excellent amino acid balance
Fish solubles (50% solids)  3      3   Excellent amino acid balance
Linseed meal                5      5   Low lysine
Meat and bone meal          5      5   Low lysine, tryptophan and
                                       methionine, good phosphorus
Peanut meal, expeller       0      5   Low lysine
Skim milk, dried           20      0   Excellent amino acid source,
                                          palatable
Sorghum gluten meal         0      5   Low lysine
Soybeans, whole, cooked    40     30   Similar to soybean meal, but
                                       may produce soft pork
Soybean meal, solvent      35     22   Similar to soybean meal,
                                          dehulled
Soybean meal, solvent,     30     20   Good amino acid balance with 
dewhulled                                  corn
Tankage (meat meal)         0      5   Low digestibility, unpalatable
Yeast, brewers dried        3      3   Source of B-vitamins
_____________________________________________________________________

1Based on an air dry basis unless otherwise noted. High  moisture
feedstuffs  must  be converted to an air dry equivalent of 88-90%
dry matter to determine energy and substitution  rates.  Complete
data on all ingredients are not available.

2When fed at no more than maximum recommended % of complete diet.
Relative   values  based  on  metabolizable  energy,  lysine  and
phosphorus content using simultaneous equations. Example:

     ME    Lysine   Phorphorus   Price
___________________________________________________________
1550X +   0.24Y +      0.25Z =   $/cwt. corn
1460X +   2.90Y +      0.60Z =   $/cwt. soybean meal (44%)
   0X +      0Y +     18.50Z =   $/cwt. dicalcium phosphate
Determine values for X, Y, and Z and multiply them times the M.E.
(kcal/lb), % lysine, and % phosphorus of feed in question and sum
the values.
3Higher levels may be fed although performance may decrease.
_____________________________________________________________________
______________________________________________________________________


List of Figures:

Figure 1: Partition of energy in nutrition.


REV 6/91 (5M)

______________________________________________

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